顶育毛蕨是金星蕨科 Thelypteridaceae毛蕨属 Cyclosorus濒危植物。1

植株高逾1米。根状茎长而横走，灰褐色，疏被褐色的披针形鳞片；叶柄深禾秆色；叶片几和叶柄等长，干后褐绿色；孢子囊群圆形，生侧脉的近先端，靠近叶边。

形态特征植株高逾1米。根状茎长而横走，粗达7毫米，灰褐色，疏被褐色的披针形鳞片。叶远生；叶柄长约55厘米，基部粗3-4毫米，疏被鳞片，向上光滑，深禾秆色；叶片几和叶柄等长，或有时稍短，宽30-36厘米，先端羽裂，急剧收缩成长尾，基部不变狭，二回羽裂；羽片约15对，几无柄，斜展，互生，彼此相距2.5-3厘米，中部以下的侧生羽片长18-25厘米，宽1.4-1.6厘米，线状披针形，先端长渐尖，基部圆楔形，对称，羽裂深1/2-1/3；裂片30-38对，斜展，长宽各约3-4毫米，三角形，短尖头，有倒三角形的缺刻分开；顶生羽片长达10厘米，中部宽1.5厘米，披针形，深羽裂，其基部裂片长达1厘米，披针形；叶脉下面清晰，侧脉每裂片7-8对，斜上，基部一对出自主脉基部，其先端交结成钝三角形网眼，并自交接点有一较长的外行小脉伸达缺刻，第二对侧脉伸达缺刻的透明膜质联线；第三对伸达缺刻以上的叶边。叶纸质，干后褐绿色，除羽轴上面有毛外，仅沿羽州下面及叶脉，有极短的柔毛和硫磺色腺体疏生。孢子囊群圆形，生侧脉的近先端，靠近叶边，下部2-3对侧脉不育，沿羽轴两侧形成宽的（约5毫米）不育空间；囊群盖小，厚膜质，棕色，无毛，宿存。2

生长环境主要生长于海拔380米的灌丛下潮湿沙土上。

分布范围主要分布于海南（感恩、昌江）、台湾（台北、台中）。广布于亚洲热带，日本、越南、泰国、缅甸、印度、马来西亚、印度尼西亚、波利尼西亚、密克罗尼西亚以及澳大利亚北部等地；非洲赞比亚也有。2

相关类别缩羽毛蕨 Cyclosorus abbreviatus Ching et Shing ex Shing et J. F. Cheng

渐尖毛蕨 Cyclosorus acuminatus (Houtt. ) Nakai

锐片毛蕨 Cyclosorus acutilobus Ching

锐尖毛蕨 Cyclosorus acutissimus Ching ex Shing et J. F. Cheng

线羽毛蕨 Cyclosorus angustipinnus (Ching) Shing

狭羽毛蕨 Cyclosorus angustus Ching

干旱毛蕨 Cyclosorus aridus (Don) Tagawa

下延毛蕨 Cyclosorus attenuatus Ching ex Shing

腺饰毛蕨 Cyclosorus aureoglandulifer Ching ex Shing

金腺毛蕨 Cyclosorus aureo-glandulosus Ching et Shing ex Ching et C. F. Zhang

百色毛蕨 Cyclosorus baiseensis Ching ex Shing

短柄毛蕨 Cyclosorus brevipes Ching et Shing

多耳毛蕨 Cyclosorus caii Ching ex Shing

三合毛蕨 Cyclosorus calvescens Ching

苍南毛蕨 Cyclosorus cangnanensis Shing et C. F. Zhang

程氏毛蕨 Cyclosorus chengii Ching ex Shing

秦氏毛蕨 Cyclosorus chingii Z. Y. Liu ex Ching et Z. Y. Liu

慈利毛蕨 Cyclosorus ciliensis Shing

棒腺毛蕨 Cyclosorus clavatus Shing

狭缩毛蕨 Cyclosorus contractus Ching ex Shing

鳞柄毛蕨 Cyclosorus crinipes (Hook. ) Ching

狭基毛蕨 Cyclosorus cuneatus Ching ex Shing

大明山毛蕨 Cyclosorus damingshanensis Ching ex Shing

光盖毛蕨 Cyclosorus decipiens Ching

德化毛蕨 Cyclosorus dehuaensis Ching et Shing

密羽毛蕨 Cyclosorus densissimus Ching ex Shing

齿牙毛蕨 Cyclosorus dentatus (Forssk.) Ching

疏羽毛蕨 Cyclosorus dissitus Ching ex Shing

独龙江毛蕨 Cyclosorus dulongjiangensis W. M. Chu

高株毛蕨 Cyclosorus elatus Ching ex Shing

广叶毛蕨 Cyclosorus ensifer (Tagawa) Shieh.

河池毛蕨 Cyclosorus euphlebius Ching

展羽毛蕨 Cyclosorus evolutus (Bedd. ) Ching

高大毛蕨 Cyclosorus excelsior Ching et Shing

国楣毛蕨 Cyclosorus fengii Ching ex Shing

平基毛蕨 Cyclosorus flaccidus Ching et Z. Y. Liu

梣叶毛蕨 Cyclosorus fraxinifolius Ching

福建毛蕨 Cyclosorus fukienensis Ching

高雄毛蕨 Cyclosorus gaoxiongensis Ching ex Shing

光叶毛蕨 Cyclosorus glabrescens Ching ex Shing

大毛蕨 Cyclosorus grandissimus Ching et Shing

粗齿毛蕨 Cyclosorus grosso-dentatus Ching ex Shing

海南毛蕨 Cyclosorus hainanensis Ching

异果毛蕨 Cyclosorus heterocarpus (Bl.) Ching

毛脚毛蕨 Cyclosorus hirtipes Shing et C. F. Zhang

毛囊毛蕨 Cyclosorus hirtisorus (C. Chr. ) Ching

河口毛蕨 Cyclosorus hokouensis Ching

学煜毛蕨 Cyclosorus houi Ching

毛蕨 Cyclosorus interruptus (Willd. ) H. Ito

闽台毛蕨 Cyclosorus jaculosus (Christ) H. Ito

景洪毛蕨 Cyclosorus jinghongensis Ching ex Shing

九龙山毛蕨 Cyclosorus jiulongshanensis Chiu et Yao ex Ching

夔州毛蕨 Cyclosorus kuizhouensis Shing

细柄毛蕨 Cyclosorus kuliangensis (Ching) Shing

贵州毛蕨 Cyclosorus kweichowensis Ching ex Shing

宽羽毛蕨 Cyclosorus latipinnus (Benth. ) Tard.-Blot

心祁毛蕨 Cyclosorus laui Ching

雷波毛蕨 Cyclosorus leipoensis Ching et H. S. Kung

龙栖山毛蕨 Cyclosorus longqishanensis Shing

阔羽毛蕨 Cyclosorus macrophyllus Ching et Z. Y. Liu

墨脱毛蕨 Cyclosorus medogensis Ching et S. K. Wu

临沧毛蕨 Cyclosorus megongensis Ching ex Shing

冕宁毛蕨 Cyclosorus mianningensis Ching ex Shing

多网眼毛蕨 Cyclosorus mollissimus Ching ex Shing

美丽毛蕨 Cyclosorus molliusculus (Wall. ex Kuhn) Ching

多囊毛蕨 Cyclosorus multisorus Ching ex Shing

南川毛蕨 Cyclosorus nanchuanensis Ching et Z. Y. Liu

南岭毛蕨 Cyclosorus nanlingensis Ching ex Shing et J. F. Cheng

南平毛蕨 Cyclosorus nanpingensis Ching

南溪毛蕨 Cyclosorus nanxiensis Ching ex Shing

黑叶毛蕨 Cyclosorus nigrescens Ching ex Shing

倒披针毛蕨 Cyclosorus oblanceolatus Shing et C. F. Zhang

峨眉毛蕨 Cyclosorus omeigensis Ching

对羽毛蕨 Cyclosorus oppositipinnus Ching et Z. Y. Liu

对生毛蕨 Cyclosorus oppositus Ching ex Shing

腺脉毛蕨 Cyclosorus opulentus (Kaulf.) Nakai

东方毛蕨 Cyclosorus orientalis Ching ex Shing

蝶状毛蕨 Cyclosorus papilio (Hope) Ching

蝶羽毛蕨 Cyclosorus papilionaceus Shing et C. F. Zhang

宽顶毛蕨 Cyclosorus paracuminatus Ching ex Shing et J. F. Cheng

曲轴毛蕨 Cyclosorus paradentatus Ching ex Shing

长尾毛蕨 Cyclosorus paralatipinnus Ching ex Shing

岳麓山毛蕨 Cyclosorus pararidus Ching ex Shing

华南毛蕨 Cyclosorus parasiticus (L.) Farwell.

小叶毛蕨 Cyclosorus parvifolius Ching

龙胜毛蕨 Cyclosorus parvilobus Ching et Shing ex Shing

少羽毛蕨 Cyclosorus paucipinnus Ching et C. F. Zhang

齿片毛蕨 Cyclosorus pauciserratus Ching et C. F. Zhang

屏山毛蕨 Cyclosorus pingshanensis Ching et H. S. Kung

无腺毛蕨 Cyclosorus procurrens (Mett. ) Ching

兰屿大叶毛蕨 Cyclosorus productus (Kaulf. ) Ching

越北毛蕨 Cyclosorus proximus Ching et C. H. Wang

假干旱毛蕨 Cyclosorus pseudoaridus Ching ex Shing

楔形毛蕨 Cyclosorus pseudocuneatus Ching ex Shing

狭叶毛蕨 Cyclosorus pumilus Ching ex Shing

泡泡毛蕨 Cyclosorus pustuliferus Ching ex Shing

矮毛蕨 Cyclosorus pygmaeus Ching et C. F. Zhang

石生毛蕨 Cyclosorus rupicola Ching

三都毛蕨 Cyclosorus sanduensis Shing et P. S. Wang

糙叶毛蕨 Cyclosorus scaberulus Ching

锯齿毛蕨 Cyclosorus serrifer Ching ex Shing

沙坪坝毛蕨 Cyclosorus shapingbaensis Ching ex Shing

石门毛蕨 Cyclosorus simenensis Shing et C. M. Zhang

同羽毛蕨 Cyclosorus simillimus Ching ex Shing

拟渐尖毛蕨 Cyclosorus sino-acuminatus Ching et Z. Y. Liu

中华齿状毛蕨 Cyclosorus sinodentatus Ching et Z. Y. Liu

疏囊毛蕨 Cyclosorus sparsisorus Ching ex Shing

狭脚毛蕨 Cyclosorus stenopes Ching et Shing ex Shing

假渐尖毛蕨 Cyclosorus subacuminatus Ching ex Shing et J. F. Cheng

短尖毛蕨 Cyclosorus subacutus Ching

坚叶毛蕨 Cyclosorus subcoriaceus Ching ex Shing

巨型毛蕨 Cyclosorus subelatus (Bak. ) Ching

万金毛蕨 Cyclosorus subnamburensis Ching ex Shing

台湾毛蕨 Cyclosorus taiwanensis (C. Chr.) H. Ito

泰宁毛蕨 Cyclosorus tarningensis Ching

河边毛蕨 Cyclosorus transitorius Ching ex Shing

截裂毛蕨 Cyclosorus truncatus (Poir.) Farwell

黄志毛蕨 Cyclosorus wangii Ching

望谟毛蕨 Cyclosorus wangmoensis Shing et P. S. Wang

温州毛蕨 Cyclosorus wenzhouensis Shing et C. F. Zhang

武陵毛蕨 Cyclosorus wulingshanensis C. M. Zhang

寻乌毛蕨 Cyclosorus xunwuensis Ching et J. F. Cheng

雁荡毛蕨 Cyclosorus yandangensis Ching et Shing

元江毛蕨 Cyclosorus yuanjiangensis Ching ex Shing

云南毛蕨 Cyclosorus yunnanensis Ching ex Shing

朝芳毛蕨 Cyclosorus zhangii Shing3

本词条内容贡献者为:

王建林 - 教授 - 兰州大学